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Tones in example sentence

Edgar, flashing with fury, dominated all the others with his clearer voice; Ashton hurled homicidal provocations at him in deep notes; Lucie uttered her shrill plaint, Arthur at one side, his modulated tones in the middle register, and the bass of the minister pealed forth like an organ, while the voices of the women repeating his words took them up in chorus delightfully.
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